Ref T12287. See D17361. That fixed a mostly-theoretical bug with crumbs named things like "MMMMMMMMMMMMMMMMMMMM", but caused a less-theoretical buggy side effect in Safari.
For now, just keep the "MMMMMMMM" crumbs around since that's the easiest/least-bad/safest fix prior to the release cut. We can fix this more broadly when we have more time to look at it.